1 00:00:53,527 --> 00:00:56,519 The sunlit waters of a shallow sea. 2 00:00:56,407 --> 00:00:59,399 Life here is rich and varied. 3 00:01:02,167 --> 00:01:08,686 Jellyfish, sea gooseberries and all kinds of larvae drift in the dappled waters. 4 00:01:09,727 --> 00:01:16,246 On the bottom, sea anemones wave among the rocks and small worms wriggle within crannies. 5 00:01:16,447 --> 00:01:20,838 Everywhere you look, there seems to be life of some kind. 6 00:01:27,047 --> 00:01:30,756 Creatures like these have a very ancient ancestry. 7 00:01:30,887 --> 00:01:34,323 They were among the first forms of life to appear on earth 8 00:01:34,247 --> 00:01:39,685 and they existed for several hundred million years before the development of fish, 9 00:01:40,007 --> 00:01:42,282 the first animals with backbones. 10 00:01:49,727 --> 00:01:54,881 But when such creatures with no bones in them die, what remains of them? 11 00:01:55,007 --> 00:02:00,445 Almost nothing. Their soft tissues simply disintegrate and dissolve in the water, 12 00:02:00,287 --> 00:02:05,042 and there's hardly anything left of them but a little slime in the mud. 13 00:02:05,567 --> 00:02:12,405 Only a minority, a few molluscs with hard shells, crustaceans like crabs with external skeletons, 14 00:02:12,287 --> 00:02:17,884 only they leave any signs of their existence after their flesh has vanished. 15 00:02:18,047 --> 00:02:20,800 So of all that multitude of creatures, 16 00:02:20,927 --> 00:02:27,366 only one or two could leave behind in the mud any evidence that they had ever existed. 17 00:02:29,087 --> 00:02:35,765 This, too, was once mud at the bottom of a sea but that was over 500 million years ago 18 00:02:36,247 --> 00:02:40,923 and now it's mudstone and high in the Canadian Rockies. 19 00:02:41,047 --> 00:02:45,484 And these rocks, too, contain the remains of the hard parts of sea animals, 20 00:02:45,367 --> 00:02:47,164 and very extraordinary animals, too. 21 00:02:47,287 --> 00:02:50,757 They're now totally extinct and we call them trilobites. 22 00:02:51,127 --> 00:02:54,722 But there's virtually nothing else but trilobites in these rocks, 23 00:02:54,967 --> 00:02:58,960 so what did the trilobites live on and, maybe, what hunted the trilobites? 24 00:02:59,287 --> 00:03:02,279 The answers to questions like those could only be guesswork 25 00:03:02,167 --> 00:03:05,477 until, that is, the year 1901. 26 00:03:14,807 --> 00:03:18,561 In that year, an American geologist, Charles Walcott, 27 00:03:18,647 --> 00:03:22,356 was exploring here in the Rocky Mountains of British Columbia, 28 00:03:22,487 --> 00:03:26,275 travelling on horseback with a train of pack mules. 29 00:03:26,327 --> 00:03:31,481 He was in his 60th year and coming towards the end of a long and distinguished career 30 00:03:31,607 --> 00:03:37,477 in which he had made a special study of the ancient fossil-bearing rocks of North America. 31 00:03:40,527 --> 00:03:43,803 When he got to this precise point on the trail, 32 00:03:44,367 --> 00:03:49,282 where this slip of loose rocks crosses it, one of his horses stumbled. 33 00:03:49,167 --> 00:03:52,796 Walcott dismounted to clear the path, and when he did so, 34 00:03:53,007 --> 00:03:57,717 he hit one of the boulders with his hammer, as he must have done ten thousand times before. 35 00:03:57,807 --> 00:03:59,877 Only this time, when the boulder fell apart 36 00:04:00,207 --> 00:04:05,361 it revealed a fossil the like of which he had never seen before in all his experience. 37 00:04:05,487 --> 00:04:10,481 To his amazement, he saw that it had its soft parts preserved: 38 00:04:10,767 --> 00:04:16,160 Tentacles, the head and a row of small legs on either side of its body. 39 00:04:16,327 --> 00:04:20,002 If he didn't do so then, he must have realised very soon afterwards 40 00:04:20,167 --> 00:04:23,682 that this was the most important discovery of his life. 41 00:04:23,527 --> 00:04:27,042 So the next season, he and his sons returned to this place 42 00:04:27,367 --> 00:04:30,677 to try and find out where that boulder had come from. 43 00:04:34,607 --> 00:04:39,601 They climbed up the rock tip, looking for fossils as they went 44 00:04:39,887 --> 00:04:47,282 and knowing that the highest level at which they found any fragments with fossils in them 45 00:04:47,087 --> 00:04:51,160 must be the place from which all the fossils were coming. 46 00:04:51,407 --> 00:04:55,161 And that proved to be just here, 47 00:04:55,247 --> 00:04:59,718 and this place has been the site of research ever since. 48 00:05:06,687 --> 00:05:12,364 A band of these shales just seven foot thick produced all the fossils. 49 00:05:12,447 --> 00:05:19,398 Walcott came here for the next eight seasons, and in that time he collected 61,000 specimens, 50 00:05:19,647 --> 00:05:24,960 and two-thirds of the species that he found proved to be new to science. 51 00:05:29,967 --> 00:05:35,963 Animals such as these, with delicate legs, with tiny gills and threadlike antennae, 52 00:05:36,127 --> 00:05:40,086 must have been living throughout the seas of this very ancient period. 53 00:05:40,447 --> 00:05:44,759 They had never been seen before because everywhere else, being soft-bodied, 54 00:05:44,767 --> 00:05:48,601 they had simply dissolved and disappeared without trace. 55 00:05:48,607 --> 00:05:52,964 Only here, for some extraordinary reason, had they been preserved, 56 00:05:53,327 --> 00:05:57,286 and preserved, what's more, in amazing detail. 57 00:05:57,167 --> 00:06:00,955 This was some kind of worm, presumably a burrower in the mud, 58 00:06:01,487 --> 00:06:04,524 with an extendible proboscis. 59 00:06:04,367 --> 00:06:08,406 And here a most remarkable find, a worm with fleshy legs, 60 00:06:08,687 --> 00:06:10,917 a possible link between true worms 61 00:06:11,087 --> 00:06:15,399 and more complex invertebrates such as centipedes and insects. 62 00:06:17,887 --> 00:06:21,038 These rocks are known as the Burgess Shales. 63 00:06:21,247 --> 00:06:25,035 How is it that one thin band of them on this particular mountainside 64 00:06:25,087 --> 00:06:29,046 preserved signs of life that are found nowhere else in the world? 65 00:06:29,407 --> 00:06:35,039 That was one of the questions Walcott and his successors spent a long time trying to answer. 66 00:06:35,167 --> 00:06:38,318 The latest group of scientists to work on the site here 67 00:06:38,527 --> 00:06:42,884 come from the Royal Ontario Museum and are led by Des Collins. 68 00:06:43,047 --> 00:06:48,246 530 million years ago, this was a muddy sea floor about 400 feet deep 69 00:06:48,327 --> 00:06:51,364 and was directly in front of a massive, sheer cliff 70 00:06:51,687 --> 00:06:55,362 that you can see in this light-coloured material here. 71 00:07:07,487 --> 00:07:11,275 The reef front rose in a sheer cliff about 300 feet high. 72 00:07:11,327 --> 00:07:16,037 You can see at the top the bedded rock, which is where the animals lived in a lagoon 73 00:07:16,127 --> 00:07:18,482 about 100 feet deep or less. 74 00:07:19,007 --> 00:07:22,522 Every so often, the mud at the top would come down in a slump, 75 00:07:22,367 --> 00:07:24,756 picking up the animals, bringing them down here, 76 00:07:25,247 --> 00:07:29,559 killing them and burying them and preserving them in the mud. 77 00:07:30,527 --> 00:07:34,759 This happened at a time when complex animals had only just appeared, 78 00:07:35,327 --> 00:07:40,355 so the ultimate ancestors of all life today must therefore be among them. 79 00:07:40,127 --> 00:07:43,278 This worm, with an internal rod running along its length, 80 00:07:43,487 --> 00:07:47,162 may be the ancestor of all backboned animals. 81 00:07:47,367 --> 00:07:50,882 And this, with five pairs of claws on its head, 82 00:07:51,207 --> 00:07:55,280 may be the creature from which scorpions and spiders have evolved, 83 00:07:55,527 --> 00:07:59,202 for it shares some of their most significant characters. 84 00:07:59,887 --> 00:08:03,675 This, with what seems a protrudable proboscis, is more of a puzzle. 85 00:08:04,167 --> 00:08:08,240 It may be related to certain kinds of living worms. 86 00:08:10,287 --> 00:08:13,916 And this bizarre creature is quite unlike anything alive today. 87 00:08:14,127 --> 00:08:18,757 It had seven pairs of legs, seven tentacles on its back, each with a mouth. 88 00:08:18,687 --> 00:08:23,363 It seems to have been one of evolution's experiments that simply didn't work very well, 89 00:08:23,487 --> 00:08:25,523 and it's left no descendants. 90 00:08:26,367 --> 00:08:28,437 By examining the best of these specimens, 91 00:08:28,767 --> 00:08:34,285 it's possible to deduce from the flattened outlines what it was like before it was squashed flat, 92 00:08:34,527 --> 00:08:37,724 and to reconstruct it in three dimensions. 93 00:08:37,887 --> 00:08:41,118 The liquid mud often penetrated the insides of the animals, 94 00:08:41,247 --> 00:08:46,605 separating each tiny organ from the other by a microscopic film of mud particles, 95 00:08:46,527 --> 00:08:49,997 and then it's possible to work out details of internal anatomy. 96 00:08:50,367 --> 00:08:57,159 This creature swam by waving flaps beneath it and it sensed its food with long feelers. 97 00:08:57,087 --> 00:09:01,683 But some specimens are so strange, it's difficult to make head or tail of them. 98 00:09:02,047 --> 00:09:07,167 This, for example, three or four inches long, looks like some kind of shrimp, 99 00:09:07,327 --> 00:09:10,524 except that none has ever been found with a head. 100 00:09:11,927 --> 00:09:17,559 And this, rather like a slice of pineapple. Could this be some kind of jellyfish? 101 00:09:17,727 --> 00:09:19,843 Though there are still some speculations, 102 00:09:19,647 --> 00:09:22,844 we now have a picture of a large and varied community. 103 00:09:23,007 --> 00:09:26,716 But if there were so many of these mud munchers and filter feeders, 104 00:09:27,327 --> 00:09:31,923 there must surely have been some hunter that preyed on them. What was that? 105 00:09:33,967 --> 00:09:37,562 That question troubled a British palaeontologist, Harry Whittington, 106 00:09:37,807 --> 00:09:40,640 as he worked on some of Walcott's specimens. 107 00:09:40,687 --> 00:09:42,359 Searching through many thousands of them, 108 00:09:42,607 --> 00:09:48,000 he found one in which that pineapple slice seemed to be attached to some other structure. 109 00:09:47,887 --> 00:09:50,924 What is more, there were several other specimens rather like it, 110 00:09:51,087 --> 00:09:54,443 including one that was not completely cleared of matrix, 111 00:09:54,607 --> 00:09:58,441 and he started very carefully to investigate it. 112 00:09:58,927 --> 00:10:01,521 ...if you think this is the underside of the body, 113 00:10:01,647 --> 00:10:05,686 to look and see, is there anything perhaps attached to the underside 114 00:10:05,487 --> 00:10:07,478 that goes down into the rock? 115 00:10:07,807 --> 00:10:10,162 And there was a little area here. 116 00:10:10,207 --> 00:10:16,077 And using a little drill, you could very delicately work along the edge here 117 00:10:16,447 --> 00:10:18,722 and remove flakes of rock. 118 00:10:18,847 --> 00:10:23,557 And gradually this thing became exposed 119 00:10:23,567 --> 00:10:30,086 and I realised, particularly when I got to this end and saw the characteristic spines on it, 120 00:10:30,287 --> 00:10:34,166 that was this thing that had been described many years before, 121 00:10:34,527 --> 00:10:38,520 anomalocaris, the strange shrimp, 122 00:10:38,567 --> 00:10:43,482 and people thought this was part of an animal and envisaged it having a little shell here, 123 00:10:44,247 --> 00:10:46,966 but no whole one had ever been found. 124 00:10:47,127 --> 00:10:51,166 Now here was one attached under the front end of this animal. 125 00:10:51,367 --> 00:10:57,203 Was that an accident? If there was one one side, there ought to be one the other. 126 00:10:57,127 --> 00:11:01,882 Indeed, there was a layer in the rock here 127 00:11:01,927 --> 00:11:05,476 and I exposed parts of that chiselling around here, 128 00:11:05,767 --> 00:11:12,081 and there exposed is part of the companion one that was attached there. 129 00:11:13,447 --> 00:11:15,483 That predator was revealed. 130 00:11:15,847 --> 00:11:21,205 The headless shrimps were its claws and the pineapple slice was its muscular mouth. 131 00:11:21,127 --> 00:11:24,119 This was the terror of the trilobites. 132 00:11:27,007 --> 00:11:29,726 So now we have an even more complete picture 133 00:11:30,207 --> 00:11:34,803 of the life that flourished on the sea floor 530 million years ago. 134 00:11:35,007 --> 00:11:39,000 We knew it must have contained the ancestors of all subsequent life, 135 00:11:38,847 --> 00:11:42,999 but now we have some idea of what they looked like. 136 00:11:45,087 --> 00:11:48,762 The next exceptionally detailed glimpse we get of the progress of life 137 00:11:48,927 --> 00:11:53,796 comes from this valley in southern Germany, near a village called Solnhofen. 138 00:11:53,647 --> 00:11:59,244 140 million years ago, that's 400 million years after the time of the Burgess Shales, 139 00:11:59,407 --> 00:12:04,276 this part of Europe lay under the sea, as we know because it's covered by limestone, 140 00:12:04,767 --> 00:12:07,964 but this valley is lined by cliffs of a special kind. 141 00:12:09,607 --> 00:12:12,519 When you examine the rock of these huge towers, 142 00:12:12,487 --> 00:12:17,038 you see that it's composed not of thin, horizontal layers of sediment, 143 00:12:17,527 --> 00:12:20,087 but of curving plates. 144 00:12:19,927 --> 00:12:25,843 And these are, in fact, the fossilised remains of sponges and other reef-building organisms 145 00:12:26,287 --> 00:12:28,642 which, growing one on top of the other, 146 00:12:28,647 --> 00:12:33,402 over centuries slowly built these huge, pillar-like reefs. 147 00:12:36,407 --> 00:12:38,602 While they were living, beneath the sea, 148 00:12:38,807 --> 00:12:41,844 they sheltered the water lying between them and the shore 149 00:12:41,687 --> 00:12:44,201 from the waves and currents of the open ocean 150 00:12:44,567 --> 00:12:47,479 and they created a shallow lagoon. 151 00:12:47,447 --> 00:12:52,475 Conditions then must have been rather like they are today in other shallow tropical lagoons 152 00:12:52,767 --> 00:12:57,204 where the water evaporates so fast that it becomes extremely salty, 153 00:12:57,567 --> 00:13:01,037 and calcium carbonate in solution starts to precipitate 154 00:13:00,927 --> 00:13:06,126 as an extremely fine, limy mud, layer upon layer. 155 00:13:06,207 --> 00:13:10,086 Over millions of years, the mud compacted and turned to stone. 156 00:13:10,527 --> 00:13:13,678 The land level rose and the sea drained away. 157 00:13:13,927 --> 00:13:19,559 The river in the Solnhofen valley eroded much of it, exposing those coral cliffs again. 158 00:13:20,647 --> 00:13:24,526 But elsewhere, the limestone remains. 159 00:13:24,887 --> 00:13:29,005 And now, as stone, those layers can be separated, where it's weathered, 160 00:13:29,207 --> 00:13:32,882 into plates almost as thin as paper. 161 00:13:33,047 --> 00:13:36,926 And where it's not weathered, it formed a magnificent building stone 162 00:13:36,847 --> 00:13:41,045 that's been worked in places like these since Roman times. 163 00:13:42,127 --> 00:13:47,121 If the rock splits into plates an inch or so thick, it can be used for roofing. 164 00:13:47,407 --> 00:13:51,559 Where it's more massive, it's cut into blocks for masonry of such quality 165 00:13:51,727 --> 00:13:55,163 that it has been used all over Germany. 166 00:13:55,527 --> 00:13:59,805 In the 19th century, a local man discovered yet another use for it. 167 00:13:59,927 --> 00:14:02,760 If you draw a picture on it with wax pencil, 168 00:14:02,807 --> 00:14:08,643 you can, using a special ink, take an almost unlimited number of copies from it. 169 00:14:08,567 --> 00:14:10,876 He called the process lithography, 170 00:14:11,447 --> 00:14:17,636 and it was widely used to print illustrations for a hundred years or more, all over Europe. 171 00:14:20,447 --> 00:14:23,962 But every now and again, when the quarrymen come to split a block, 172 00:14:24,287 --> 00:14:28,485 they open it and find that there's an illustration already printed within it, 173 00:14:28,607 --> 00:14:32,486 an illustration that's 140 million years old, 174 00:14:32,767 --> 00:14:34,758 like this one 175 00:14:36,127 --> 00:14:37,640 A fish. 176 00:14:38,047 --> 00:14:44,520 Fossils are not abundant, for few animals could live in this inhospitable, near-sterile lagoon. 177 00:14:44,767 --> 00:14:49,238 These fish were swept into it, past the reefs, by sudden storms. 178 00:14:49,087 --> 00:14:52,921 But though there are few of them, their preservation is near-perfect, 179 00:14:53,407 --> 00:14:56,763 for the storm that carried them in also stirred up the mud 180 00:14:56,767 --> 00:15:01,966 and, as they died, that settled back and covered them like a shroud. 181 00:15:03,047 --> 00:15:06,835 A fish with grinding teeth, like a parrot-fish of today, 182 00:15:07,367 --> 00:15:12,999 which presumably fed in the same way, pulverising coral to extract the little polyps. 183 00:15:14,807 --> 00:15:16,718 A kind of garfish. 184 00:15:25,407 --> 00:15:27,238 A bottom-living ray, 185 00:15:27,327 --> 00:15:32,196 a species that seems to have survived almost unaltered until today. 186 00:15:39,247 --> 00:15:44,275 This lobster, too, had ancestors that strayed into the Solnhofen lagoon. 187 00:15:50,767 --> 00:15:53,725 Crustaceans, with their hard external skeletons, 188 00:15:53,847 --> 00:15:57,078 make excellent subjects for preservation under these conditions. 189 00:16:00,607 --> 00:16:03,565 And backboned animals, too, with their skeletons of bone, 190 00:16:03,487 --> 00:16:05,523 are also beautifully preserved. 191 00:16:05,887 --> 00:16:10,085 A turtle, its flesh gone but its bones very clear. 192 00:16:10,847 --> 00:16:15,967 Another reptile, with an amazingly long body and an even longer tail. 193 00:16:16,127 --> 00:16:19,119 Yet, because of the infrequency of the currents in the lagoon, 194 00:16:19,487 --> 00:16:24,481 all its bones remained perfectly positioned while the flesh decayed around them. 195 00:16:24,767 --> 00:16:26,917 And because of that extreme stillness, 196 00:16:26,687 --> 00:16:33,957 even a jellyfish settling gently on the limy mud has left the delicate impress of its soft body. 197 00:16:38,487 --> 00:16:42,196 Tracks on the lagoon floor have also remained clear. 198 00:16:42,327 --> 00:16:46,036 This one was obviously made by an animal with several long legs. 199 00:16:46,167 --> 00:16:50,399 The scratches between the footprints suggest that it also had a spiky tail. 200 00:16:50,487 --> 00:16:54,685 And it seems to have been lost, for it's wandering aimlessly about. 201 00:16:54,807 --> 00:17:01,326 We're looking at the story of one small death that took place 140 million years ago, 202 00:17:01,527 --> 00:17:05,600 for here's the body, complete with that trailing tail. 203 00:17:07,447 --> 00:17:11,440 It's a horseshoe crab, virtually identical with the horseshoe crabs 204 00:17:11,767 --> 00:17:15,999 that still today swim in the seas off North America. 205 00:17:16,087 --> 00:17:18,237 Other tracks are not so easily interpreted. 206 00:17:19,927 --> 00:17:25,638 What about, for example, this extremely rare but rather mysterious mark? 207 00:17:25,767 --> 00:17:28,839 Is it perhaps some kind of worm? 208 00:17:29,127 --> 00:17:32,756 Well, it seems to have been caused by an empty ammonite shell 209 00:17:32,967 --> 00:17:35,845 that fell down into the mud, making a dent, 210 00:17:35,887 --> 00:17:40,199 and then, carried by a current, rolled until it came to the mouth again, 211 00:17:40,687 --> 00:17:46,876 which caused it to leap up, fall again, and then roll again. 212 00:17:48,367 --> 00:17:53,805 Insects flying over the lagoon sometimes flopped into the water and sank. 213 00:17:54,127 --> 00:17:56,118 A dragonfly. 214 00:17:58,527 --> 00:18:01,280 A winged grasshopper. 215 00:18:01,727 --> 00:18:05,197 And a much bigger flying animal, a pterosaur, 216 00:18:05,087 --> 00:18:08,921 the membrane of its skinny wings plainly visible. 217 00:18:15,247 --> 00:18:17,283 But there's one animal in particular, 218 00:18:17,647 --> 00:18:22,846 whose remains drifted down through the salty water and settled on the mud of the lagoon, 219 00:18:22,927 --> 00:18:27,045 that has made the name Solnhofen famous worldwide. 220 00:18:27,247 --> 00:18:32,196 Its remains are so excessively rare, and so important to science, 221 00:18:32,527 --> 00:18:36,998 that they've been called "the most valuable fossils in the world". 222 00:18:36,847 --> 00:18:40,556 The most perfect of them, which is kept here in East Berlin, 223 00:18:41,167 --> 00:18:46,366 is so valuable that it's kept locked away in a safe, away from public view, 224 00:18:46,447 --> 00:18:49,519 and exhibited only as a replica. 225 00:18:49,327 --> 00:18:54,117 This is a rare privilege to see the real thing. 226 00:19:00,207 --> 00:19:05,281 Archaeopteryx, a creature that represents a link between reptiles and birds. 227 00:19:09,327 --> 00:19:13,764 It's birdlike because it's covered not by fur but by feathers. 228 00:19:14,127 --> 00:19:19,121 Their intricate structure, each with a central quill and barbs coming off it on either side, 229 00:19:19,207 --> 00:19:21,198 can be clearly seen. 230 00:19:21,127 --> 00:19:24,722 They are virtually identical to modern feathers. 231 00:19:26,887 --> 00:19:29,321 Its wings are modified front legs, 232 00:19:29,767 --> 00:19:33,123 but not as greatly altered as the wings of modern birds, 233 00:19:33,127 --> 00:19:36,483 for three of their five toes still have claws at their tips, 234 00:19:36,607 --> 00:19:39,883 projecting from the front edge of the wings. 235 00:19:44,807 --> 00:19:50,040 The feet had backward-pointing big toes, which gave the animal a firm grip on a branch, 236 00:19:50,567 --> 00:19:52,956 and that, too, is characteristic of birds. 237 00:19:52,967 --> 00:19:55,765 The head, however, is not at all birdlike. 238 00:19:55,887 --> 00:20:00,438 It had no beak, but a bony, reptilian jaw lined with teeth. 239 00:20:04,087 --> 00:20:10,276 And here, reptilian and bird characteristics combined in one feature, the tail. 240 00:20:10,327 --> 00:20:13,319 No bird has bones in its tail like this, 241 00:20:13,687 --> 00:20:17,475 and while reptiles do, none of them has feathers on it. 242 00:20:17,567 --> 00:20:21,719 In recent years, the question was raised as to whether these really were feathers, 243 00:20:21,887 --> 00:20:25,084 but the closest examination of another specimen in London 244 00:20:25,247 --> 00:20:28,284 has proved conclusively that they certainly are. 245 00:20:29,087 --> 00:20:33,205 So, thanks to the limy burial shrouds of Solnhofen 246 00:20:33,407 --> 00:20:35,398 we can make a detailed reconstruction 247 00:20:35,807 --> 00:20:40,039 of this key creature in the history of the evolution of life. 248 00:20:45,887 --> 00:20:49,357 But reptiles were still, at this time, the dominant animals. 249 00:20:49,247 --> 00:20:53,240 Gigantic seagoing crocodiles like this one roamed the seas. 250 00:20:53,567 --> 00:20:57,355 Huge pterosaurs bigger than any eagle soared through the air. 251 00:20:57,407 --> 00:21:00,524 And on land, there were the dinosaurs. 252 00:21:00,767 --> 00:21:04,680 The reason we know so much about dinosaurs is that many were very big, 253 00:21:05,087 --> 00:21:08,636 with tough bones that could survive being washed down by the rivers 254 00:21:08,447 --> 00:21:11,280 and buried in the deposits of the delta. 255 00:21:11,807 --> 00:21:16,483 But 67 million years ago, the last of the dinosaurs died. 256 00:21:16,607 --> 00:21:19,644 For some time, the land was comparatively underpopulated, 257 00:21:19,487 --> 00:21:22,160 but then the early mammals began to spread. 258 00:21:22,367 --> 00:21:25,165 But they were not big creatures with tough bones 259 00:21:25,247 --> 00:21:29,798 but small animals with delicate skeletons that were easily destroyed. 260 00:21:30,047 --> 00:21:33,437 Neither did they live in a lagoon where there were regular deposits, 261 00:21:33,887 --> 00:21:37,243 but on land, where there was virtually none. 262 00:21:37,247 --> 00:21:41,320 So they were comparatively poor candidates for fossilisation 263 00:21:41,567 --> 00:21:45,162 and we knew very little about them until, at last, 264 00:21:45,407 --> 00:21:49,446 another of these extraordinary fossil sites was discovered. 265 00:21:50,687 --> 00:21:55,317 At Messel, near Frankfurt, in Germany, there's a deposit of shales so rich in oil 266 00:21:55,687 --> 00:22:00,078 that quite spontaneously, they catch fire and burn underground. 267 00:22:00,007 --> 00:22:03,079 The oil comes from the tissues of animals and plants 268 00:22:03,247 --> 00:22:06,922 living in the lake that lay here 48 million years ago. 269 00:22:07,087 --> 00:22:10,079 The shales were once worked commercially for their oil, 270 00:22:10,527 --> 00:22:13,917 but they contain much more valuable things than that. 271 00:22:15,687 --> 00:22:21,045 48 million years is a comparatively short time for mud to turn to rock. 272 00:22:21,327 --> 00:22:26,321 Solnhofen limestone is three times as old, the Burgess Shales over ten times, 273 00:22:26,607 --> 00:22:30,316 and these shales are still soft and moist. 274 00:22:30,407 --> 00:22:33,399 The excavators have spotted the remains of a fish, 275 00:22:33,287 --> 00:22:38,805 but it mustn't be fully exposed to the air because if it dries out, it'll disintegrate, 276 00:22:39,047 --> 00:22:43,438 so the slab is carefully cut out and taken to a laboratory. 277 00:22:46,487 --> 00:22:53,438 There it's kept moist while the compressed mud is delicately scraped from the flank of the fish. 278 00:23:07,687 --> 00:23:13,637 Once that side has been cleaned as far as possible, liquid resin is poured over it. 279 00:23:28,087 --> 00:23:31,204 When that sets, it's perfectly transparent, 280 00:23:31,127 --> 00:23:35,837 so now specimens treated in this way can be worked on from the other side. 281 00:23:43,367 --> 00:23:46,677 Eventually, virtually all the mud can be removed, 282 00:23:46,767 --> 00:23:52,205 and the fragile bones are held firmly in a block of transparent plastic. 283 00:23:55,487 --> 00:23:59,924 This is one of the early mammals for which Messel is now famous, 284 00:23:59,807 --> 00:24:04,403 a tiny horse that was no bigger than a spaniel. 285 00:24:12,967 --> 00:24:16,642 Gerhard Storch, from the Senckenberg Museum in Frankfurt, 286 00:24:16,807 --> 00:24:19,401 is one of the team working on these finds. 287 00:24:19,567 --> 00:24:25,836 Unlike recent horses, they possess four digits in the forefeet, 288 00:24:25,887 --> 00:24:28,720 one, two, three, four. 289 00:24:28,767 --> 00:24:33,602 - Both legs are there, aren't they? - Yeah, there are both legs, side by side. 290 00:24:34,047 --> 00:24:38,757 - And perfect little hooves. - And perfect little hooves, yeah. 291 00:24:38,807 --> 00:24:43,483 The grinding teeth are not tall and high in the jaw like those of a modern horse, 292 00:24:43,607 --> 00:24:47,998 but low, indicating that the animal ate soft leaves and fruit, 293 00:24:47,927 --> 00:24:51,556 and that's confirmed by examining the stomach. 294 00:24:52,047 --> 00:24:58,600 With a scanning microscope, we can investigate the last diet of these horses. 295 00:24:58,727 --> 00:25:05,917 In this specimen, the gut contained soft leaves from tropical plants. 296 00:25:05,927 --> 00:25:12,605 - So it was living, what, in the woodlands? - It was living in a dense, tropical forest. 297 00:25:12,807 --> 00:25:17,198 And there are even stranger things in the muds of Messel than spaniel-sized horses. 298 00:25:17,327 --> 00:25:22,959 This is a mammal that belongs to a very archaic group close to insectivores 299 00:25:23,087 --> 00:25:25,726 and this group became extinct. 300 00:25:26,567 --> 00:25:30,560 The forelimbs are very small, very reduced. 301 00:25:30,407 --> 00:25:32,875 The hind limbs are long, elongated. 302 00:25:33,287 --> 00:25:37,200 The tail is very long. It's a world record for mammals. 303 00:25:37,287 --> 00:25:42,042 It consists of about 50 single vertebrae. 304 00:25:42,087 --> 00:25:45,397 And all of these body proportions tell us 305 00:25:45,447 --> 00:25:49,360 that this animal was moving on its hind limbs, 306 00:25:49,847 --> 00:25:55,126 but not in a way that is familiar to us from kangaroos or jerboas, 307 00:25:55,087 --> 00:26:00,878 running fast and very manoeuvrable on the hind limbs but with alternating steps. 308 00:26:01,327 --> 00:26:03,887 (ATTENBOROUGH) So really there's no equivalent alive today? 309 00:26:03,727 --> 00:26:06,639 (STORCH) There is no analogue today. 310 00:26:09,887 --> 00:26:13,516 There is also, believe it or not, an ancestor of hedgehogs. 311 00:26:13,607 --> 00:26:16,280 The texture of bone on the front of the skull 312 00:26:16,487 --> 00:26:20,116 shows that the flesh there was particularly thick with blood vessels, 313 00:26:20,287 --> 00:26:24,360 which suggests that there was some kind of gland or shield on its forehead. 314 00:26:25,287 --> 00:26:28,085 And there are true birds, many kinds, 315 00:26:28,167 --> 00:26:34,003 some so excellently preserved that almost the only detail you can't make out is their colour. 316 00:26:34,007 --> 00:26:39,559 And not only birds, but bats, and several kinds of them as well. 317 00:26:42,967 --> 00:26:47,483 But why should so many flying animals have fallen dead into the lake? 318 00:26:47,967 --> 00:26:54,122 Something must have happened to these bats while they were hunting on the wing, 319 00:26:54,207 --> 00:26:59,884 and my idea is that there were poisonous gases on the Messel lake 320 00:26:59,807 --> 00:27:04,403 and a bat which went down drinking came to such a gas bubble 321 00:27:04,607 --> 00:27:06,837 and fell down to the water's surface. 322 00:27:09,887 --> 00:27:13,721 So, because of those freakish conditions 40 million years ago, 323 00:27:14,207 --> 00:27:16,516 the vanished lake of Messel now yields evidence 324 00:27:16,607 --> 00:27:20,680 about not only the animals that swam in its waters and lived in its forests, 325 00:27:20,927 --> 00:27:23,441 but even those that populated the skies. 326 00:27:26,767 --> 00:27:28,758 Los Angeles, in America. 327 00:27:28,687 --> 00:27:32,282 Hardly world-famous for its fossils, but it should be, 328 00:27:32,527 --> 00:27:35,360 for in the heart of this most modern of cities 329 00:27:35,887 --> 00:27:41,598 is a site that gives another wholly exceptional picture of a vanished world. 330 00:27:44,527 --> 00:27:46,518 40,000 years ago, 331 00:27:46,927 --> 00:27:51,318 this was the appearance of the land on which hotels and freeways now stand. 332 00:27:51,247 --> 00:27:55,638 Firm evidence for every single detail in this most detailed painting 333 00:27:56,047 --> 00:28:01,440 comes from a small park close by one of the city's main avenues, La Brea. 334 00:28:03,207 --> 00:28:06,324 In one corner of it, through the harmless grass, 335 00:28:06,567 --> 00:28:12,039 oozes a substance that kills - la brea, tar. 336 00:28:16,567 --> 00:28:20,719 It wells up from the ground here to form these black pools. 337 00:28:20,887 --> 00:28:23,355 When it rains, water lies on top of it, 338 00:28:23,767 --> 00:28:26,645 and it looks like a place where you might get a drink. 339 00:28:26,647 --> 00:28:31,084 But any animal that came here to do so would be lucky to escape alive. 340 00:28:30,967 --> 00:28:36,963 Feet sink into the tar, feathers get entangled in it, and the animal is fatally trapped. 341 00:28:37,207 --> 00:28:40,756 That's been happening for 40,000 years and more, 342 00:28:41,167 --> 00:28:44,284 and it's still happening today. 343 00:28:53,887 --> 00:28:58,119 Tar, like oil, is derived from the bodies of animals and plants 344 00:28:58,207 --> 00:29:00,198 that accumulated in swamps. 345 00:29:00,127 --> 00:29:03,722 The sand that was deposited on top of them squeezed their remains 346 00:29:03,967 --> 00:29:07,198 so that droplets of oil were expelled from their tissues. 347 00:29:07,327 --> 00:29:11,605 That accumulated in basins within the texture of the porous sandstone 348 00:29:12,127 --> 00:29:17,485 and then, where there is a fault, this substance is forced up to the surface. 349 00:29:19,327 --> 00:29:22,717 The earlier flows of tar, containing the most ancient animals, 350 00:29:22,687 --> 00:29:25,042 have now been covered by later flows, 351 00:29:25,087 --> 00:29:28,921 so to reach them, you have to dig down into the tar pit, 352 00:29:29,407 --> 00:29:32,763 and excavations which started back at the beginning of this century 353 00:29:32,767 --> 00:29:35,600 are now being carried on some 30 feet down, 354 00:29:35,647 --> 00:29:39,640 while the tar still rises around the excavating platform. 355 00:29:40,567 --> 00:29:43,320 The work is supervised by trained scientists, 356 00:29:43,447 --> 00:29:47,565 but most of the team is made up of local volunteers. 357 00:29:47,727 --> 00:29:51,766 (MAN) 0K. This is ready to go. Got the bag? (W0MAN) Yes, I do. 358 00:29:52,167 --> 00:29:55,125 (MAN) That's a sabre-toothed cat femur. (W0MAN) Right or left? 359 00:29:55,527 --> 00:29:57,597 (MAN) That's a right. 360 00:29:57,927 --> 00:30:00,725 (W0MAN) And without his epiphysis. (MAN) Right. 361 00:30:00,487 --> 00:30:05,117 (MAN) What you got over there, Jerry? (JERRY) I've got this ulna uncovered here. 362 00:30:05,447 --> 00:30:10,202 - What's it lying on top of? - It's lying right across a sabre-tooth cat skull. 363 00:30:10,247 --> 00:30:14,923 The finds have been put on display in a museum recently built on the site, 364 00:30:15,407 --> 00:30:17,443 and very spectacular they are. 365 00:30:28,127 --> 00:30:31,199 In addition to this magnificent imperial mammoth, 366 00:30:31,487 --> 00:30:34,923 the biggest of all the prehistoric elephants that lived in North America, 367 00:30:35,327 --> 00:30:40,845 there were extinct horses and camels which grazed on these plains. 368 00:30:43,927 --> 00:30:47,124 Huge, long-horned bison. 369 00:30:49,287 --> 00:30:53,360 There were over 20 species of eagles and falcons. 370 00:30:54,327 --> 00:30:58,081 Ground sloths the size of small elephants. 371 00:30:58,167 --> 00:31:01,204 These huge animals are now totally extinct. 372 00:31:01,527 --> 00:31:04,963 They browsed the trees and, like the rest of these plant-eaters, 373 00:31:04,887 --> 00:31:08,357 only rarely and accidentally strayed into the tar pits. 374 00:31:08,727 --> 00:31:11,639 But once an animal like this was mired and stuck, 375 00:31:11,607 --> 00:31:15,805 it made the pits a positive attraction to packs of wolves. 376 00:31:18,847 --> 00:31:22,442 These dire wolves were about the same size as living wolves 377 00:31:22,487 --> 00:31:24,318 but with more massive heads. 378 00:31:24,407 --> 00:31:28,195 Struggling, trapped animals were obviously something they couldn't resist, 379 00:31:28,247 --> 00:31:32,604 for wolves, in fact, are the commonest of all the victims of the tar pits. 380 00:31:34,887 --> 00:31:38,084 The most frequently trapped grass-eaters were the bison, 381 00:31:38,247 --> 00:31:40,681 so there were probably big herds of them, 382 00:31:40,647 --> 00:31:44,686 but, again, the pits contain more bones of the animal that preyed on them, 383 00:31:44,967 --> 00:31:47,083 the American lion. 384 00:31:55,287 --> 00:31:58,802 The females were about the same size as African lions, 385 00:31:59,127 --> 00:32:02,597 but the males were 25% bigger. 386 00:32:09,207 --> 00:32:13,678 And there was an even more impressive cat, the sabre-tooth. 387 00:32:14,967 --> 00:32:19,245 At one time it was thought that these extraordinary teeth were daggers for stabbing, 388 00:32:19,447 --> 00:32:24,282 but now it's believed that they were used to slit open the belly of the prey. 389 00:32:24,247 --> 00:32:28,081 You might wonder how the animal managed even to close its jaws, 390 00:32:28,407 --> 00:32:32,480 and I asked a scientist at the museum, George Jefferson, to explain. 391 00:32:32,727 --> 00:32:37,323 I think we can illustrate that best with the specimen here that was preserved closed. 392 00:32:37,447 --> 00:32:41,360 That will give us an idea what that mouth looked like. 393 00:32:41,287 --> 00:32:48,477 As you can see, the incisors actually interlaced, allowing the jaw to fully close. 394 00:32:48,847 --> 00:32:51,042 I'm surprised how much space there is 395 00:32:51,247 --> 00:32:54,796 between those huge sabre teeth and the lower jaw. 396 00:32:55,007 --> 00:33:01,037 That gap is the same gap as between the meat-slicing teeth on the side of the face. 397 00:33:00,767 --> 00:33:05,079 What that meant is, the animal would disengage the incisors, 398 00:33:05,567 --> 00:33:08,843 drop the jaw down, move it slightly sideways, 399 00:33:08,927 --> 00:33:11,725 and guide the slicing blades here 400 00:33:11,767 --> 00:33:17,000 by running the inside of this flange against the canine tooth. 401 00:33:17,047 --> 00:33:20,244 - Is that a new discovery? - It is. In fact, 402 00:33:20,607 --> 00:33:25,078 we didn't know this gap was that way until we found this specimen. 403 00:33:27,327 --> 00:33:32,640 As well as big animals, the tar preserved a whole range of organisms that lived here. 404 00:33:32,607 --> 00:33:35,075 Even pollen grains can be obtained from it, 405 00:33:35,487 --> 00:33:39,719 and it's they that have enabled the artist who painted the reconstructed landscape 406 00:33:39,807 --> 00:33:45,120 to show the correct species of flowering bush, Californian sage. 407 00:33:48,927 --> 00:33:52,203 The museum's laboratory is surrounded by glass windows, 408 00:33:52,287 --> 00:33:56,519 so that those working on the finds become exhibits themselves. 409 00:33:57,087 --> 00:33:59,840 I think this rib's going in like that, 410 00:34:00,327 --> 00:34:03,444 and if you look at the top, the way it angles round, 411 00:34:03,687 --> 00:34:08,238 - I think there's probably a little edge. - So it's going back under. 412 00:34:08,007 --> 00:34:12,603 Once you've discovered what animals were present, why go on digging up more? 413 00:34:12,887 --> 00:34:16,562 Well, the museum has over 200 jaws of bison. 414 00:34:16,927 --> 00:34:20,966 As George Jefferson explained, their sheer number gives new information. 415 00:34:20,767 --> 00:34:22,758 With these three specimens here, 416 00:34:23,167 --> 00:34:27,285 we can see an animal three years old, two years old and one year old. 417 00:34:28,927 --> 00:34:33,205 We can tell that from the eruption stage of the teeth. 418 00:34:33,327 --> 00:34:38,162 Here we see this tooth being pushed out - this animal is three years old. 419 00:34:38,127 --> 00:34:41,005 We don't have specimens representing intermediate ages 420 00:34:41,007 --> 00:34:45,603 between three years old and two years old, nor two and one year old. 421 00:34:46,087 --> 00:34:50,763 That tells us that these animals are coming here periodically at a certain time of year. 422 00:34:50,887 --> 00:34:55,915 We can determine that season by comparing the stage of wear on these teeth 423 00:34:56,167 --> 00:34:59,637 with modern bison who have a very restricted calving period. 424 00:34:59,527 --> 00:35:01,438 They're here in the late springtime. 425 00:35:01,927 --> 00:35:03,519 - So they're migrating? - Yes. 426 00:35:05,087 --> 00:35:09,126 The multitudes of jaws, therefore, prove that once in California, 427 00:35:08,927 --> 00:35:12,283 great herds of bison made long migratory journeys, 428 00:35:12,767 --> 00:35:17,443 and that every year, after the rains, when the grass began to sprout on the Hollywood hills, 429 00:35:17,567 --> 00:35:20,843 there was great carnage around the tar pits. 430 00:35:31,447 --> 00:35:35,918 The sheer abundance of the dire-wolf skulls also yields information. 431 00:35:36,167 --> 00:35:41,480 They are not, of course, all the same, and the differences are not all due to age. 432 00:35:41,327 --> 00:35:47,038 The lumps and distortions that are apparent, compared to the smooth forehead on this animal, 433 00:35:47,647 --> 00:35:50,878 indicate an infection in the frontal sinuses of the forehead, 434 00:35:51,447 --> 00:35:55,486 probably as the result of being kicked in the face, 435 00:35:55,527 --> 00:35:58,325 maybe by a bison or a camel. 436 00:35:58,647 --> 00:36:02,196 This animal was obviously going after its prey and getting injured. 437 00:36:02,967 --> 00:36:05,561 We also see injuries in the sabre cats. 438 00:36:06,807 --> 00:36:09,526 We see chronic injuries to the back. 439 00:36:11,127 --> 00:36:15,245 Here we have three lumbar or back vertebrae 440 00:36:15,327 --> 00:36:19,286 that have been fused together by a mass of bony tissue. 441 00:36:19,527 --> 00:36:24,442 This bony tissue grows along the ligaments and muscles where they're injured and stretched 442 00:36:24,327 --> 00:36:26,966 while this animal is lunging for its prey. 443 00:36:30,967 --> 00:36:32,958 In this hip... 444 00:36:34,407 --> 00:36:38,480 ...we have a fairly normal hip socket here, 445 00:36:38,247 --> 00:36:43,640 but on this side you can see a lot of knobbly bone, 446 00:36:44,007 --> 00:36:46,805 a distortion and break, 447 00:36:46,887 --> 00:36:50,641 there are flanges of extra bone in here. 448 00:36:50,727 --> 00:36:55,039 This animal was obviously in a lot of pain while this healing was going on. 449 00:36:55,527 --> 00:36:59,202 We think it may have been butted by a bison, hit very hard, 450 00:36:59,367 --> 00:37:01,927 or even, possibly, by a mammoth elephant. 451 00:37:01,767 --> 00:37:04,076 - So he's really a cripple. - Yes. 452 00:37:04,647 --> 00:37:07,081 It's astounding it lived as long as it did. 453 00:37:07,047 --> 00:37:10,517 Some researchers believe this is evidence 454 00:37:10,527 --> 00:37:16,477 that the injured and infirm were being tolerated within the population and possibly cared for. 455 00:37:16,767 --> 00:37:19,964 - So social behaviour among the sabre cats? - Social behaviour. 456 00:37:22,167 --> 00:37:25,239 It seems almost miraculous that any remains of animals 457 00:37:25,527 --> 00:37:30,555 could survive for tens of thousands of years, let alone hundreds of millions. 458 00:37:30,607 --> 00:37:35,840 In exceptional circumstances, like the tar pits of La Brea or the volcanic lake of Messel, 459 00:37:36,367 --> 00:37:41,885 the hot lagoons of Solnhofen and the muddy submarine avalanches of the Burgess Shales, 460 00:37:42,007 --> 00:37:45,238 they leave clues that tell us not only about their appearance, 461 00:37:45,367 --> 00:37:51,681 but their detailed internal anatomy, their daily habits, even their social life. 462 00:37:51,607 --> 00:37:54,997 Fossils have been forming ever since life appeared on this planet 463 00:37:55,447 --> 00:37:58,007 and lie in the earth all around us. 464 00:37:57,847 --> 00:38:03,524 They provide us with irrefutable and wonderful evidence of what existed before we did. 465 00:38:05,527 --> 00:38:09,679 Sometimes the latest high-tech apparatus is needed to reveal it, 466 00:38:09,847 --> 00:38:13,362 sometimes it needs nothing more than a simple blow from a hammer 467 00:38:13,207 --> 00:38:15,482 and logical, clear-minded thought. 468 00:38:16,087 --> 00:38:17,884 But slowly, piece by piece, 469 00:38:18,007 --> 00:38:21,716 we're putting together the history of the long procession of life 470 00:38:21,847 --> 00:38:25,078 that preceded mankind's appearance upon this planet, 471 00:38:25,207 --> 00:38:28,279 and of which, indeed, we are a part.